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By D. I. Edwards

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2 Streptomyces spp. 2 2 The -group numbers indicate differences in the shape and structural disposition of the macrolide ring and the subgroup letter indicates differences in the disposition of the sugars attached to it. 52 ANTIMICROBIAL DRUG ACTION The biosynthesis of erythromycin differs from that of the fused ring antibiotics discussed previously in that the poly ketone structure of the macrolide ring derives from seven propionate residues rather than acetate. Methymycin also derives from the condensation of propionate units whereas carbomycin derives from acetate condensation.

Some of the steps shown have not been fully elucidated. 7. -AA side chain characteristic of cephalosporin C. -AA moiety has the D- configuration in penicillin N and the L- configuration in isopenicillin N. 7 Biogenetic origin of cephalosporins. 8 Structure of penicillin N. 4 Gramicidin S The biosynthesis of this antibiotic is better known than that of the tyrocidins which it resembles, since it has been synthesised in a cell-free system which enables biosynthetic analysis of the steps to be made with relative ease.

H. Hiatt, J. D. Watson and J. A. Winston eds), Cold Spring Harbor Publications, New York, pp. 1473-82 Schubert, J. (1970). Formation of cell growth inhibition by chemicals and environmental agents. f. gen. , 64, 37-40 Zahner, H. and Maas, W. K. (1972). Biology of Antibiotics. 1 INTRODUCTION A knowledge of how antimicrobial drugs are biosynthesised is essential for the production and purification of such drugs and provides a means whereby new drugs can be synthesised chemically. All the major antibiotics in use today are produced by bacteria or fungi with the exception of small antibiotic molecules such as chloramphenicol which can be synthesised chemically.

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